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Life Science Dictionary

Phloem

Botany

Say it: FLOH-em

Phloem is the living vascular tissue of plants that carries sugars and other organic nutrients from leaves to the parts that use or store them.

Also: bast

Definition

What is phloem?

Phloem is the plant vascular tissue that distributes the products of photosynthesis, mainly the sugar sucrose, along with amino acids, hormones and signaling molecules. Its conducting cells, sieve-tube elements, are joined end to end into sieve tubes, with porous end walls called sieve plates. Unlike the water-conducting cells of xylem, they are alive at maturity, although they have lost their nucleus and most organelles. Each one depends on a neighboring companion cell, which carries out much of its metabolism and is linked to it by many tiny channels.

Phloem moves sap from sources, such as mature leaves, to sinks, such as roots, fruits, seeds and growing shoots. The accepted explanation is the pressure-flow model. Sucrose is actively loaded into sieve tubes at the source, which costs energy; water then enters by osmosis from nearby xylem, building pressure that pushes the sap along. At the sink, sucrose is unloaded and water leaves, keeping pressure low at that end. Because sinks change as a plant grows, phloem can carry material up or down.

The name comes from the Greek for bark, because in woody stems phloem lies just inside the bark.

Word parts

Where does the word “phloem” come from?

phloe-
barkGreek phloos

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